COMBAT

Tower Guide — The Full Roster

Every tower in the game with live stats, plus how stars, branches, strains and mastery mutate them.

The roster

Every tower is an organism with a combat class, a firing pattern and a role on the line. This table is generated from the live game data, so it is always current with the shipped build:

Tower	Class	Cost	Range	Damage	Cadence	Splash	Role
Needle Spitter	Kinetic	100	700	3	0.4s	—	Very long range — peppers from afar, stacking Fracture (armor shred).
Bone Mortar	Caustic	170	960	8	2.1s	60	Full-map artillery. Slow, arcing AoE; leaves Corrosion pools (DoT + slow).
Flesh Reactor	Vital	120	90	0	0s	—	Generates biomass; overcharges NEARBY towers (+dmg, +rate).
Parasite Nest	Viral	130	280	6	0.8s	—	Medium range — place forward. Seeds Infection: ramping armor-ignoring DoT.
Neural Obelisk	Neural	165	360	5	1.1s	—	Wide control field — slows + makes enemies take more from ALL sources.
Carrion Maw	Kinetic	140	170	26	1.3s	—	Short-reach jaws — CLEAVES everything inside them. The last line eats well.
Amber Chrysalis	Caustic	155	430	18	3.4s	—	Encases the BIGGEST threat in amber — frozen solid, briefly out of the fight.
Wailing Effigy	Neural	150	300	0	2.6s	—	A shrieking totem — panics attackers into FLEEING the Hive. Buys time, not kills.
Resonance Coil	Resonant	150	340	26	0.5s	—	Locks a beam onto ONE target; damage RAMPS the longer it holds — melts elites & bosses.
Marrow Siphon	Vital	175	300	22	0.4s	—	Splits a draining WEB across the wounded — GORGES fatter the more the lane dies, siphoning the swarm into Biomass.
Blight Sower	Caustic	130	300	46	2.8s	88	Seeds the approach with ARMED spore mines — pre-placed detonations that corrode everything they catch.
Synapse Spire	Neural	150	210	30	1.1s	—	Casts a synaptic ARC that leaps victim to victim — the tighter the pack, the longer the lightning.
Undertow Polyp	Kinetic	140	165	5	1.6s	—	A rooted undertow that DRAGS the wave back up-lane with every pulse — progress itself becomes the resource you deny.
Spitfang Turret	Kinetic	90	380	12	0.6s	—	A reliable mid-range autocannon — brisk, punchy single-target fire. The dependable damage pick.
Rift Anchor	Neural	175	190	20	1.4s	—	A premium void anchor — a psionic singularity that DRAGS the wave to its heart and crushes it. The deepest unlock.

Four evolution layers

A tower kind grows along four independent axes. Stars (permanent, bought with Tower Shards in the Lab) climb an authored ★-path of stat and behavior upgrades. Branches (in-run, per instance) lock a tower into one of two mutually exclusive mutations that re-author how it fires. Strain (in-run) is the evolution layer fed by consuming duplicate organisms from the Biomorph Conveyor. Mastery (permanent XP from kills) unlocks ranks, lore and cosmetic evolution.

The practical order for a new player: place towers → learn their branches → push stars on your favorites → let mastery accumulate on its own.

Reading a tower’s role

Cost and range tell you where it lives; cadence and splash tell you what it kills. Cheap + long range = your opener. High splash = swarm control. Beam or debuff organs multiply everything AROUND them — they never carry a line alone. Artillery has a deadzone: enemies closer than its minimum range cannot be targeted, so screen it with short-range brawlers.